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Guiyu oneiros , on display at the Paleozoological Museum of China .

Guiyu oneiros

An early bony fish fossil from the Silurian period of China.

Photo: Sun Jiao (Interaccoonale), CC BY 4.0, via Wikimedia Commons

Guiyu oneiros is a fossil species of bony fish discovered in Yunnan, China. It dates to the late Silurian period, approximately 425 million years ago. The find helps scientists understand how ray-finned and lobe-finned fishes evolved.

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  • Discovered in Yunnan, China, dating to 425 million years ago.
  • Holotype specimen is 26 cm long; estimated living length is 30 cm.
  • Possessed placoderm-like spines on its fins and dermal pelvic bones.
  • Described in 2009 by Zhu Min based on an articulated fossil.
  • Part of the Xiaoxiang fauna, including placoderms and acanthodians.

What it is

Guiyu oneiros represents one of the earliest known articulated bony fish fossils. The specimen measures about 26 centimetres in length with a depth of 11 centimetres. Scientists estimate the living animal reached roughly 30 centimetres long. Its body featured a streamlined shape, while the head accounted for nearly a quarter of its total length. The creature possessed primitive traits typical of placoderms, such as large spines on its pectoral, pelvic, and dorsal fins. Unlike modern bony fish, its pelvis included dermal bones alongside endochondral bone. Three median dorsal plates sat behind the skull roof, with the third plate supporting the first dorsal fin spine. These bones were covered in ganoine, a hard tissue also found in basal ray-finned fishes.

Where it lives

Fossils of this species originate from the Kuanti Formation near Qujing in Yunnan province, China. The remains were recovered from late Silurian marine strata, specifically dated to the late Ludlow age using conodonts as index fossils. This location is part of the Xiaoxiang fauna, a rich collection of marine ecosystem fossils. The site also yields remains of galeaspids, placoderms, and acanthodians. The geological context places the species in a diverse marine environment from approximately 425 million years ago.

How it lives

The classification of Guiyu has shifted over time. Initially described as a basal lobe-finned fish with mixed features, it is now often viewed as a stem-group osteichthyan. This placement suggests it lies outside the main split between ray-finned and lobe-finned fishes. Some analyses recover it as a sister group to crown-osteichthyans. These primitive characteristics indicate that the divergence of major bony fish lineages occurred by the late Silurian period. The presence of placoderm-like features highlights the complex evolutionary path of early jawed vertebrates.

Guiyu oneiros
A drawing made by EveryFish from what this page says. It is a diagram, not a picture of the real fish.

Fish and people

The species was formally described in 2009 by Chinese palaeontologist Zhu Min and colleagues. Their description relied on a near-complete articulated specimen. The generic name Guiyu translates from Mandarin as 'ghost fish'. The specific name oneiros derives from the Greek word for 'dream'. This naming reflects the elusive nature of such ancient fossils. The discovery provides critical data for understanding the early diversification of bony fishes during the Palaeozoic era.

Questions people ask about Guiyu oneiros

What does the name Guiyu mean?

The name Guiyu comes from the Mandarin term guǐyú, which translates to 'ghost fish'. The specific epithet oneiros is derived from the Greek word for 'dream', reflecting the elusive nature of these ancient fossils.

How old are Guiyu fossils?

The fossils date to the late Silurian period, approximately 425 million years ago. They were found in the late Ludlow-aged Kuanti Formation in Yunnan, China, identified using conodonts as index fossils.

Is Guiyu a modern fish?

No, Guiyu is an extinct fossil species. It is considered a basal osteichthyan, likely a stem-group member lying outside the main split between ray-finned and lobe-finned fishes that exist today.

Good reading and sources

Where this page comes from

The facts on this page come from the Wikipedia article "Guiyu oneiros", written by its contributors and used under the CC BY-SA 4.0 licence, and from its Wikidata record (CC0). EveryFish rewrote the text in its own words, with AI assistance, and compares every page with the article to make sure no passage is copied. It may be reused under the same CC BY-SA 4.0 licence with a link back.

It may contain mistakes, and figures change over time. Nothing here is investment or buying advice. If something is wrong, tell us. Last refreshed 2026-11-15.

Photo credits on this page

The small pictures above come from Wikimedia Commons under free licences. Some are cropped to fit.

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